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Geranium cuneatum subsp. hololeucum

(A.Gray) Carlquist & Bissing (1976)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Geraniales Family: Geraniaceae Genus: Geranium

Description

Key Characters:

Growth Form: Compact, erect, many-branched shrubs 3–10 dm tall.

Stems: Stems and branches reddish to black, covered with persistent stipules, densely leafy only toward apex.

Roots:

Leaves: Leaves simple. Alternate. Blades oblong–cuneate, (1.5–)2.5–5 cm long, (0.5–)0.7–1.2(–1.4) cm wide. Apex truncate to sometimes rounded. Surfaces densely grayish to sometimes silky strigose on both surfaces. Margins entire except apex (3)4–8-toothed. Usually 3–8-veined. Petioles 0.3–2 cm long. Stipules subulate, ca. 5–6 mm long, strigillose.

Flowers: Flowers 7–30 in compound cymes, forming large corymbose inflorescences that project beyond the leaves. Flowers bisexual (perfect), actinomorphic; pedicels 0.5–1.6(–2) cm long. Calyx of 5 sepals; sepals narrowly lanceolate, 5–9 mm long, densely silky strigose to sparsely villous or merely ciliate, apex mucronate. Petals white or cream, sometimes with purple veins and sometimes purple at base, narrowly obovate, 9–15 mm long. Stamens (5)10; filaments ± connate at base, those alternate with the petals longer than others and with basal glands, staminal filaments usually 9–12 mm long, glands pubescent; anthers dithecal, opening by longitudinal slits. Ovary superior or nearly so, 5-celled, carpels connate around a central column to form a compound ovary with as many cells, fertile portion a lobed ring at base of stylar column, placentation axile; ovules 2 per cell, anatropous to campylotropous, usually pendulous; styles 3–5, 5–6 mm long at anthesis, elongating to 15–28 mm long in fruit, essentially beakless, sometimes narrowed below apex, branches filiform, reflexed; stigmas slender and dry, rarely capitate.

Fruit: Septicidal and elastically dehiscent capsules separating into as many segments as carpels; a portion of the style splitting off from remainder of stylar column and forming an awn that recurves upward from the persistent central column; usually remaining attached to apex; sometimes the awn also becomes spirally coiled; awn usually hygroscopic; carpel bodies ca. 2–4 mm long; sparsely to densely pubescent. Seeds 1 per cell; dark reddish brown; oblong–obovoid; ca. 1.7–1.9 mm long; the surface inconspicuously reticulate.

Ploidy:

Habitat: Occasional to forming a dominant part of the vegetation in subalpine shrubland and forest; alpine shrubland; and as a pioneer in shrubland on lava in fog swept areas.

Elevation Range: 1,850–3,050 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Biotropica 8: 258 (1976)

Other References

Wagner et al. 1990:730 (H); Aedo 2023:579 (DESCR, KEY)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing NE of Halekohaku, Mauna Kea arid lava rock and ledges PRESERVED_SPECIMEN Degener, O. 19962 Hawaii BISH 9/1/1949
2 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing ne of halekohaku, mauna kea PRESERVED_SPECIMEN degener, o. murashige, t.; kerr, m. collector number: 19962 BISH 9/1/1949
3 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Hosmer, R.S. 51 Hawaii BISH 11/1/1909
4 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Rock, J.F.C. 25614 Hawaii BISH 12/1/1955
5 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing 0.1 mi Hilo side, Puu Huluhulu, Saddle Rd. red pahoehoe flow PRESERVED_SPECIMEN Stemmermann, R.L. 1161 Hawaii BISH 8/21/1976
6 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing saddle road near puu huluhulu `a`a lava flow PRESERVED_SPECIMEN degener, o. greenwell, a. collector number: 19979 Hawaii BISH 8/20/1949
7 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, Wailuku River scrub PRESERVED_SPECIMEN St. John, H. 26959 Hawaii BISH 6/6/1980
8 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, west slope PRESERVED_SPECIMEN Skottsberg, C.J.F. 702 Hawaii BISH 9/28/1922
9 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing 0.1 mi. hilo side, puu huluhulu, saddle rd. red pahoehoe flow PRESERVED_SPECIMEN stemmermann, r.l. collector number: 1161 Hawaii BISH 8/21/1976
10 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing e of humuula, below kalaieha puu forest PRESERVED_SPECIMEN neal, m.c. collector number: 688 Oahu BISH 8/7/1935
11 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN rock, j.f.c. collector number: 25614 Hawaii BISH 12/1/1955
12 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing hale pohaku, mauna kea Locality, mauka of buildings, below winding road. Habitat: open mamane forest. PRESERVED_SPECIMEN davis, j. collector number: 610 Hawaii BISH 10/6/1981
13 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauka of kulani prison disturbed wet lowland forest; Metrosideros, Acacia, Dicranopteris PRESERVED_SPECIMEN degener, o. greenwell, a. collector number: 21862 Oahu BISH 2/15/1952
14 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea cinder, boulders, with Styphelia PRESERVED_SPECIMEN Nagata, K.M. 1713 Hawaii BISH 7/17/1978
15 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Kahulu PRESERVED_SPECIMEN Russ, G.W. s.n. Hawaii BISH 7/1/1927
16 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, above Hale Pohaku steep volcanic ash; with Styphelia douglasii, Gnaphalium, Vaccinium PRESERVED_SPECIMEN Iltis, H.H. 204 Hawaii BISH 6/23/1967
17 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Humuula Trail to summit of Mauna Kea PRESERVED_SPECIMEN Forbes, C.N. 843 H Hawaii BISH 6/14/1915
18 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Puu Hualalai with Styphelia tameiameiae, Fragaria chiloensis, Sisyrinchium acre, Osteomeles anthyllidifolia; volcanic cinders PRESERVED_SPECIMEN Carlquist, S. 2098 Hawaii BISH 7/30/1966
19 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing kalaeha, mauna kea PRESERVED_SPECIMEN hillebrand, w.b. collector number: s.n. Hawaii BISH 1862-08-08
20 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Saddle Road near Puu Huluhulu `a`a lava flow PRESERVED_SPECIMEN Degener, O. 19979 Hawaii BISH 8/20/1949
21 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Kalaeha, Mauna Kea PRESERVED_SPECIMEN Hillebrand, W.B. s.n. Hawaii BISH 2/4/1905
22 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hale Pohaku Browsed by herbavores. PRESERVED_SPECIMEN Corn, C. s.n. Hawaii BISH 1/17/1980
23 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN rock, j.f.c. collector number: 8700 Hawaii BISH 8/8/1910
24 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea On precipitous slopes & ledges PRESERVED_SPECIMEN hosmer, r.s. collector number: 51 Hawaii BISH 11/1/1909
25 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hale Pohaku, Mauna Kea Locality, mauka of buildings, below winding road. Habitat: open mamane forest. PRESERVED_SPECIMEN Davis, J. 610 Hawaii BISH 10/6/1981
26 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, west slope PRESERVED_SPECIMEN skottsberg, c.j.f. collector number: 702 Hawaii BISH 9/28/1922
27 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing E Mauna Kea PRESERVED_SPECIMEN Skottsberg, C.J.F. 1968 Hawaii BISH 9/27/1926
28 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Rock, J.F.C. 8700 Hawaii BISH 8/1/1910
29 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, Wailu PRESERVED_SPECIMEN Forbes, C.N. 471.H Hawaii BISH 8/1/1911
30 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing E of Humuula, below Kalaieha Puu on pahoehoe PRESERVED_SPECIMEN Neal, M.C. 688 Hawaii BISH 8/7/1935
31 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Humuula Trail to summit of Mauna Kea PRESERVED_SPECIMEN Forbes, C.N. 843.H Hawaii BISH 6/14/1915
32 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, wailuku river PRESERVED_SPECIMEN forbes, c.n. collector number: 884.h Hawaii BISH 6/18/1915
33 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, above hale pohaku steep volcanic ash; with Styphelia douglasii, Gnaphalium, Vaccinium PRESERVED_SPECIMEN iltis, h.h. iltis, c.m. collector number: 204 Hawaii BISH 6/23/1967
34 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing e mauna kea PRESERVED_SPECIMEN skottsberg, c.j.f. collector number: 1968 Hawaii BISH 9/27/1926
35 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauka of Kulani Prison On partly shrubby `a`a lava flow PRESERVED_SPECIMEN Degener, O. 21862 Hawaii BISH 2/15/1952
36 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing hale pohaku, mauna kea, hamakua district Growing beneath dead mamane in open mamane parkland. PRESERVED_SPECIMEN cuddihy, l.w. collector number: 887 Hawaii BISH 10/6/1981
37 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Laumaia below tree line PRESERVED_SPECIMEN Neal, M.C. 863 Hawaii BISH 8/18/1935
38 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing above Mauna Kea-Humuula [Humuula trail PRESERVED_SPECIMEN Hartt, C.E. 756 Hawaii BISH 8/9/1955
39 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, wailuku river PRESERVED_SPECIMEN forbes, c.n. collector number: 885.h Oahu BISH 6/18/1915
40 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing laumaia above trees PRESERVED_SPECIMEN neal, m.c. hartt, c. collector number: 859 Hawaii BISH 8/18/1935
41 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauka of Kulani Prison On partly shrubby `a`a lava flow; common. PRESERVED_SPECIMEN Degener, O. 21862 Hawaii BISH 2/15/1952
42 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing kau, anipeahi, s mauna loa, 1-1.5 km e of lava flow dry ohia forest, with native trees, shrubs, grass; on pahoehoe PRESERVED_SPECIMEN warshauer, f.r. mceldowney, p.h. collector number: frw 2291 Hawaii BISH 6/12/1979
43 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Fosberg, F.R. s.n. Hawaii BISH 12/5/1936
44 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing humuula trail to summit of mauna kea PRESERVED_SPECIMEN forbes, c.n. collector number: 843.h Hawaii BISH 6/14/1915
45 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, wailuku river scrub PRESERVED_SPECIMEN st.john, h. collector number: 26959 Hawaii BISH 6/6/1980
46 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Puu Kanakaleonui area; Mauna Kea. Sophora Sub-Alpine Dry Forest. PRESERVED_SPECIMEN K. R. Wood 2605 Hawaii PTBG 6/3/1993
47 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hawa?» PRESERVED_SPECIMEN Degener, O. (Coll. G) G 8/20/1949
48 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hilo Forest Reserve, Saddle Rd at 35.4km Dry montane forest PRESERVED_SPECIMEN C. R. Annable 3750 NY 1/14/1998
49 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing 21 mi W of Hilo on Saddle Road. PRESERVED_SPECIMEN James S. Miller|Mary Merello Miller 7927 Missouri Botanical Garden 8/5/1992
50 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing 21 mi w of hilo on saddle road. PRESERVED_SPECIMEN james s. miller|mary merello miller 7927 MO 8/5/1992
51 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing kulani rd 8 mi above prison camp Cultivated PRESERVED_SPECIMEN f. r. fosberg 39309 NY 1/5/1958
52 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea; Puu Kanakaleonui, near campsite for Akiapola'au bird study; with Sophora chrysophylla subalpine dry forest. PRESERVED_SPECIMEN K. R. Wood 652 Hawaii PTBG 3/9/1991
53 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing maui. PRESERVED_SPECIMEN wilkes explor. exped. s.n. Maui US 1838-08-08
54 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN r. hosmer s.n. Oahu US
55 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing along saddle road near puu huluhulu PRESERVED_SPECIMEN o. degener & a. b. greenwell 19979 Maui US 8/20/1949
56 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing hawaii, mauna kea, mauka of halepohaku PRESERVED_SPECIMEN o. degener & i. degener 35314r Hawaii US 7/20/1967
57 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing source of wailuku river, mauna kea PRESERVED_SPECIMEN c. n. forbes 2539/885.h Kauai US 6/18/1915
58 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, silversword exclosure, edge of gulch at headwaters of Wiluku Asso. with Coprosma montana, Styphelia douglasii, Dubautia arborea, Vaccinium peleanum, Silene struthioloides, Asplenium trichomanes, and Pellaea ternifolia. PRESERVED_SPECIMEN Davis, J. 921 Hawaii BISH 9/21/1982
59 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing slopes of mauna kea (waiki) PRESERVED_SPECIMEN c. n. forbes 466.h Hawaii US
60 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing slopes of mauna kea (waiki) PRESERVED_SPECIMEN c. n. forbes 471.h Hawaii US
61 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea-north side PRESERVED_SPECIMEN a. s. hitchcock 14286 Hawaii US 8/22/1916
62 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Faurie, A. 852 Hawaii BISH 7/1/1909
63 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Slopes of Mauna Kea, Waiki PRESERVED_SPECIMEN Forbes, C.N. 466.H Hawaii BISH 8/1/1911
64 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Saddle Rd., 23.5 miles from Hilo lava PRESERVED_SPECIMEN Degener, O. 20208 Hawaii BISH 8/26/1949
65 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hale Pohaku, Mauna Kea, Hamakua District Growing beneath dead mamane in open mamane parkland. PRESERVED_SPECIMEN Cuddihy, L.W. Collector Number: 887 Hawaii BISH 10/6/1981
66 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing hilo forest reserve, saddle rd at 35.4km PRESERVED_SPECIMEN c. r. annable 3750 BRU 1/14/1998
67 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Maui. PRESERVED_SPECIMEN Wilkes Explor. Exped. s.n. Maui US 1838-00-00
68 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Aina Hou PRESERVED_SPECIMEN Forbes, C.N. 831.H Hawaii BISH 6/12/1915
69 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing s slope of mauna kea volcano, along rd above state park cabins (halepohaku) PRESERVED_SPECIMEN f. r. fosberg h-204 Hawaii US 6/23/1967
70 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, silversword exclosure, edge of gulch at headwaters of Wiluku Asso. with Coprosma montana, Styphelia douglasii, Dubautia arborea, Vaccinium peleanum, Silene struthioloides, Asplenium trichomanes, and Pellaea ternifolia. PRESERVED_SPECIMEN Davis, J. Collector Number: 921 Hawaii BISH 9/21/1982
71 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing 0.2 mi. up from turnoff to hale pohaku, ~28 mi. up saddle rd. from hilo, then 6.4 mi. n toward mauna kea PRESERVED_SPECIMEN stuessy, t.f. collector number: 6151 Hawaii BISH 5/30/1982
72 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, source of wailuku river PRESERVED_SPECIMEN forbes, c.n. collector number: 883.h Hawaii BISH 6/18/1915
73 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing northeast of halepohaku, mauna kea PRESERVED_SPECIMEN o. degener, t. murashige & kerr, -- 19962 Hawaii US 9/1/1949
74 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, wailu PRESERVED_SPECIMEN forbes, c.n. collector number: 471.h Hawaii BISH 8/8/1911
75 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hamakua, Hale Pohaku, Mauna Kea, ~5 mi N of jct. with Saddle Road with Sophora chrysophylla, Gnaphalium sandwicensium, Senecio sylvaticus PRESERVED_SPECIMEN Gustafson, R. 1095 Hawaii BISH 7/13/1978
76 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Kalaelia PRESERVED_SPECIMEN Hillebrand, W.B. s.n. Hawaii BISH
77 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Aina Hou PRESERVED_SPECIMEN Forbes, C.N. 828 H Hawaii BISH 6/12/1915
78 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN nagata, k.m. kimura, b. collector number: 1713 Maui BISH 7/17/1978
79 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN rock, j.f.c. collector number: 8345 Hawaii BISH 5/1/1910
80 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Where valley narrows into a cliff-sided ravine; on various rocks and boulders PRESERVED_SPECIMEN wilkes expedition s.n. Maui F
81 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Hosmer, R.S. s.n. Hawaii BISH 8/1/1911
82 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauka of kulani prison On waste ground with both parental species. PRESERVED_SPECIMEN degener, o. greenwell, a. collector number: 21862 Oahu BISH 2/15/1952
83 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing aina hou At base of damp cliff face.. PRESERVED_SPECIMEN forbes, c.n. collector number: 831.h Maui BISH 6/12/1915
84 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing hale pohaku Epiphytic on Halimeda opuntia. PRESERVED_SPECIMEN corn, c. collector number: s.n. Molokai BISH 1/17/1980
85 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing aina hou PRESERVED_SPECIMEN forbes, c.n. collector number: 828 h BISH 6/12/1915
86 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN hosmer, r.s. Hawaii BISH 8/8/1911
87 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Aina Hou PRESERVED_SPECIMEN Forbes, C.N. 829.H Hawaii BISH 6/12/1915
88 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing above mauna kea-humuula [humuula trail PRESERVED_SPECIMEN hartt, c.e. collector number: 756 Hawaii BISH 8/9/1955
89 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing aina hou PRESERVED_SPECIMEN forbes, c.n. collector number: 826 h Oahu BISH 6/12/1915
90 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing n hilo, humuula, wailuku river PRESERVED_SPECIMEN herbst, d.r. collector number: 6148 Oahu BISH 7/15/1978
91 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hale Pohaku Browsed by herbavores. PRESERVED_SPECIMEN Corn, C. Collector Number: s.n. Hawaii BISH 1/17/1980
92 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, source of Wailuku River PRESERVED_SPECIMEN Forbes, C.N. 883.H Hawaii BISH 6/18/1915
93 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing N Hilo, Humuula, Wailuku River cliff face in silversword enclosure PRESERVED_SPECIMEN Herbst, D.R. 6148 Hawaii BISH 7/15/1978
94 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing 0.2 mi up from turnoff to Hale Pohaku, ~28 mi up Saddle Rd. from Hilo, then 6.4 mi N toward Mauna Kea PRESERVED_SPECIMEN Stuessy, T.F. 6151 Hawaii BISH 5/30/1982
95 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, Wailuku River PRESERVED_SPECIMEN Forbes, C.N. 884.H Hawaii BISH 6/18/1915
96 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing kahulu PRESERVED_SPECIMEN russ, g.w. Hawaii BISH 7/1/1927
97 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing humuula trail to summit of mauna kea PRESERVED_SPECIMEN forbes, c.n. collector number: 843 h Hawaii BISH 6/14/1915
98 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hale Pohaku, Mauna Kea, Hamakua District Growing beneath dead mamane in open mamane parkland. PRESERVED_SPECIMEN Cuddihy, L.W. 887 Hawaii BISH 10/6/1981
99 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Kau, Anipeahi, S Mauna Loa, 1-1.5 km E of lava flow dry ohia forest, with native trees, shrubs, grass; on pahoehoe PRESERVED_SPECIMEN Warshauer, F.R. FRW 2291 Hawaii BISH 6/12/1979
100 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing aina hou PRESERVED_SPECIMEN forbes, c.n. collector number: 829.h Hawaii BISH 6/12/1915
101 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Hale Pohaku, Mauna Kea Locality, mauka of buildings, below winding road. Habitat: open mamane forest. PRESERVED_SPECIMEN Davis, J. Collector Number: 610 Hawaii BISH 10/6/1981
102 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea, Wailuku River PRESERVED_SPECIMEN Forbes, C.N. 885.H Hawaii BISH 6/18/1915
103 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing slopes of mauna kea, waiki PRESERVED_SPECIMEN forbes, c.n. collector number: 466.h Hawaii BISH 8/8/1911
104 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing kalaelia PRESERVED_SPECIMEN hillebrand, w.b. Hawaii BISH
105 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing hamakua, hale pohaku, mauna kea, ~5 mi. n of jct. with saddle road with Sophora chrysophylla, Gnaphalium sandwicensium, Senecio sylvaticus PRESERVED_SPECIMEN gustafson, r. collector number: 1095 Hawaii BISH 7/13/1978
106 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea PRESERVED_SPECIMEN fosberg, f.r. collector number: s.n. Hawaii BISH 12/5/1936
107 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea, silversword exclosure, edge of gulch at headwaters of wiluku Asso. with Coprosma montana, Styphelia douglasii, Dubautia arborea, Vaccinium peleanum, Silene struthioloides, Asplenium trichomanes, and Pellaea ternifolia. PRESERVED_SPECIMEN davis, j. collector number: 921 Hawaii BISH 9/21/1982
108 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Aina Hou PRESERVED_SPECIMEN Forbes, C.N. 826 H Hawaii BISH 6/12/1915
109 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauka of Halepohaku, Mauna Kea PRESERVED_SPECIMEN Degener, O. 35314 Hawaii BISH 7/20/1967
110 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing puu hualalai In hot, boulder-strewn region not far from sea with Lipochaeta degeneri. PRESERVED_SPECIMEN carlquist, s. collector number: 2098 Molokai BISH 7/30/1966
111 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Laumaia above trees PRESERVED_SPECIMEN Neal, M.C. 859 Hawaii BISH 8/18/1935
112 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing Mauna Kea PRESERVED_SPECIMEN Rock, J.F.C. 8345 Hawaii BISH 5/1/1910
113 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing mauna kea Vegetation on plots stripmined for bauxite ore in 1958-60. Base ore, no overburden or soil treatment. PRESERVED_SPECIMEN faurie, a. collector number: 852 Kauai BISH 7/1/1909
114 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing saddle rd., 23.5 miles from hilo lava PRESERVED_SPECIMEN degener, o. murashige; kerr collector number: 20208 Hawaii BISH 8/26/1949
115 Geranium cuneatum subsp. hololeucum (A.Gray) Carlquist & Bissing laumaia below tree line PRESERVED_SPECIMEN neal, m.c. hartt, c. collector number: 863 Hawaii BISH 8/18/1935
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